Fedexia striegeli: A New, Exquisitely Preserved 300 MYA Trematopid

A team of researchers from Carnegie Museum of Natural History has described a new genus and species of carnivorous amphibian from western Pennsylvania. The fossil skull, found in 2004 near Pittsburgh International Airport, was recovered from rocks deposited approximately 300 million years ago during the Late Pennsylvanian Period. Named Fedexia striegeli, it is one of only a very few relatively large amphibian fossils to display evidence of a predominantly terrestrial (land-based) life history so early in geologic time. The rocks where Fedexia was found are nearly 20 million years older than the localities of its fossil relatives, suggesting that the expansion and diversification of this group occurred much earlier than had been recognized previously. The full paper will be released today in Annals of Carnegie Museum, Volume 78, Number 4, 15 March 2010.

Fedexia was described on the basis of a remarkably well-preserved fossil skull. Unlike similar discoveries, the five-inch-long (11.5 cm) fossil skull remained three-dimensional over time because it was never crushed by rocks that were deposited above it. Fedexia striegeli was named for FedEx, the corporation that owns the land on which the fossil was found, and for amateur discoverer Mr. Adam Striegel, who originally found the specimen on a geology field trip while a senior at the University of Pittsburgh.

Fedexia represents an extinct group of amphibians called Trematopidae that lived about 70 million years before the first dinosaurs appeared. Unlike almost all other Pennsylvanian Period amphibians, which did not often venture out of the water, this rare, diverse group lived mostly on land, returning to the water perhaps only to mate or lay eggs. The trematopids also provide evidence of the earliest vertebrate life in North America adapted to a mostly terrestrial existence. Their success may have been a result of a long-term, global trend toward drier, warmer conditions that reached its climax near the end of the Pennsylvanian Period.

Soooo...does this one count as a reptilomorph or not? Is there a good, modern text on Carboniferous-Permian nonamniote vertebrate evolution out there?