First North American occurrence of the toothed pteranodontoid pterosaur Cimoliopterus
A new pterosaur species, Cimoliopterus dunni, sp. nov., is described based on a partial rostrum from the upper Cenomanian Britton Formation in the Eagle Ford Group of north-central Texas. The holotype preserves alveoli for a minimum of 26 upper teeth and bears a thin premaxillary crest that begins above the fourth pair of alveoli. The rostrum, characterized by a slight lateral flare, lacks the pronounced lateral expansion found in ornithocheirids and anhanguerids. The tip of the snout is small and blunt, and the anterior face of the rostrum is oriented posteroventrally, forming an approximately 45° angle with the anterior portion of the palate. The anteroventrally oriented first pair of alveoli is directed more ventrally than anteriorly. Phylogenetic analysis suggests that Cimoliopterus dunni and Cimoliopterus cuvieri, from the Cenomanian Grey Chalk Subgroup in Kent, England, are basal pteranodontoids that are closely related to Aetodactylus halli from the middle Cenomanian Tarrant Formation of north Texas. Cimoliopterus dunni marks only the second known occurrence of Cimoliopterus, extending its geographic range from Europe to North America. In conjunction with Coloborhynchus wadleighi from the upper Albian Pawpaw Formation, Cimoliopterus dunni provides unambiguous evidence of biogeographic linkages between the pterosaur faunas of North America and Europe in the middle Cretaceous.