Paleolithic Papers #13

Genus Homo:

Brain size and tooth size were not linked during human evolution.

Stone tools can be examined for the types of plants they were used on.

Asian populations are genetically contiguous with their stone age predecessors.

Did the first humans arrive in North America much earlier than previously thought?

Modern Humans (Homo sapiens):

Tibet may have been occupied by modern humans by the end of the Pleistocene.

Paleolithic cave art started out as public art and became private over time.

The environment of Shanxi, China during the late Pleistocene.

Human remains of the Mesolithic of South Africa.

Foraging patterns have persisted in New Guinea since the terminal Pleistocene despite the advent of farming.

Neandertals (Homo neanderthalensis):

The underrated genius of Neandertals.

Neandertals get praised and examined in the NY Times.

Neandertals were rock collectors.

Examining the molars of Neandertals.

Reconstructing the diet of Neandertals at El Sidrón, Spain.

How did Neandetals spread to the Greek Island of Naxos?  Boats?

Homo naledi:

Examining the ribs and vertebra of H. naledi.

Genus Australopithecus:

A. sediba:

A sediba was adapted to tree climbing as well as bipedal walking.

Dietary Ecology of Pleistocene Quaternary Camels

Dietary ecology of Pleistocene camelids: Influences of climate, environment, and sympatric taxa

Authors:

Yann et al

Abstract:

Wild members of Camelidae live in some of the most arid environments, including North Africa, Arabia, the Gobi Desert of China and Mongolia and high elevation environments in the Andes Mountains. A better understanding of the paleoecology of the three most abundant Pleistocene camelids (Camelops, Hemiauchenia, and Palaeolama) may clarify modern adaptations to arid environments. Mammalian tooth enamel δ13C values were used to compare diets of co-occurring species in California, Texas, and Florida and δ18O values were used to investigate climate. Carbon isotope analysis suggests Camelops was likely an opportunistic browser that consumed both C3 and C4 browse/CAM plants, potentially consuming C4 browse (e.g., saltbush). Hemiauchenia had an opportunistic and highly generalized diet, while Palaeolama was a specialized forest browser. Stable oxygen isotopes and aridity index values suggest that Ingleside was warmer than McKittrick Brea, but there are no significant differences in aridity between the two sites. Co-occurrence data from the Paleobiology Database suggest that Palaeolama was restricted to forested environments as it occurred with two browsers, Tapirus and Odocoileus, at 90.5% of all sites. Camelops and Hemiauchenia both co-occurred with a broader range of taxa, further suggesting these camelids lived in diverse habitats. The generalized diet of Hemiauchenia, the likely ancestor of modern South American camelids, allowed for the adaptations of extant Lama and Vicugna to survive in the arid environments of the Andes Mountains. Collectively, these data clarify the dietary ecology of extinct camelids and provide insight into the potential importance of generalist diets for increased resilience to changing environments and/or climates.

Pleistocene Giant Sloths Found in Brazil

New Pleistocene remains of megalonychid ground sloths (Xenarthra: Pilosa) from the intertropical Brazilian region

Authors:

De Iuliisa et al

Abstract:

The Pleistocene fossil sloth Australonyx aquae De Iuliis, Cartelle, and Pujos, 2009 (Mammalia, Xenarthra, Megalonychidae) was described from the intertropical region of Brazil. However, its mandible was not known and only cursory descriptions of the ear ossicles were included. The mandible was subsequently recognized among the remains originally collected from the type locality, and belongs to the holotype individual. As a particularly important skeletal element for specific recognition, it requires description to complement our understanding of this species. The ossicles, usually poorly represented in the fossil record, require further description to allow differentiation from those of other sloths. Comparisons of the mandible and ossicles are conducted with homologous elements of the contemporaneous and sympatric Ahytherium aureum Cartelle, De Iuliis, and Pujos, 2008, the only other megalonychid sloth known from intertropical Brazil, and reinforce the distinction between these two species detailed in their initial descriptions. Comparisons with other sloths (e.g., Acratocnus, Megalonyx, Neocnus) also reveal differences with Au. aquae in such features as form and size of the caniniform tooth, angular process, and mandibular condyle. Differences among the malleus and incus of Au. aquae and several species of other sloth clades reveal clade level distinctions among Megatheriidae, Nothrotheriidae, and Megalonychidae. A well-preserved skull from the Brazilian state of Rondônia is noted as probably belonging to Au. aquae. This skull cannot be assigned formally to this species because it is not deposited in a recognized institution, but it does extend considerably the known range of the species.

Evidence of a Supernova Found in Pleistocene Microfossil Record

Time-resolved 2-million-year-old supernova activity discovered in Earth’s microfossil record

Authors:

Ludwig et al

Abstract:

Massive stars (M≳10M⊙), which terminate their evolution as core-collapse supernovae, are theoretically predicted to eject >10−5M⊙ of the radioisotope 60Fe (half-life 2.61 Ma). If such an event occurs sufficiently close to our solar system, traces of the supernova debris could be deposited on Earth. Herein, we report a time-resolved 60Fe signal residing, at least partially, in a biogenic reservoir. Using accelerator mass spectrometry, this signal was found through the direct detection of live 60Fe atoms contained within secondary iron oxides, among which are magnetofossils, the fossilized chains of magnetite crystals produced by magnetotactic bacteria. The magnetofossils were chemically extracted from two Pacific Ocean sediment drill cores. Our results show that the 60Fe signal onset occurs around 2.6 Ma to 2.8 Ma, near the lower Pleistocene boundary, terminates around 1.7 Ma, and peaks at about 2.2 Ma.

Ground Sloths Spread Into North America Multiple Times

The manus of Mylodon darwinii Owen (Tardigrada, Mylodontidae) and its phylogenetic implications

Authors:

Haro et al

Abstract:

The first nearly complete and articulated manus of the ground sloth Mylodon darwinii, from the upper Pleistocene of Argentina, is described. It shares similarities with Mylodonopsis ibseni from Brazil, including a cuneiform with a distinct pisiform facet, an obliquely concave ulnar facet, and a prominent distolateral process, as well as a gracile metacarpal III. It shares a flattened pisiform with Glossotherium robustum. The trapezoid is unique in the obliquely elongate proportions of its dorsal surface. Shapes of the articular facets indicate different functions in digits II and III, with the former having a greater range of motion and the latter greater stability at the joints. Clear arboreal or fossorial adaptations are absent. A phylogenetic analysis recovered M. darwinii as closely related to M. ibseni and agrees with larger phylogenetic analyses of sloths based on craniomandibular evidence. Our data support more than two mylodontine dispersal events to North America.

Modern Europeans are Descended From Anatolian Farmers, not Paleolithic Europeans

For most of the last 45,000 years Europe was inhabited solely by hunter-gatherers. About 8,500 years ago a new form of subsistence - farming - started to spread across the continent from modern-day Turkey, reaching central Europe by 7,500 years ago and Britain by 6,100 years ago. This new subsistence strategy led to profound changes in society, including greater population density, new diseases, and poorer health. Such was the impact of farming on how we live that scientists have debated for more than 100 years how it was spread across Europe. Many believed that farming was spread as an idea to European hunter-gatherers but without a major migration of farmers themselves.

This week, an international research team led by paleogeneticists of Johannes Gutenberg University Mainz (JGU) publishes a study in the journal Proceedings of the National Academy of Sciences of the United States of America showing that early farmers from across Europe have an almost unbroken trail of ancestry leading back to the Aegean. The scientists analyzed the DNA of early farmer skeletons from Greece and Turkey. According to the study, the Neolithic settlers from northern Greece and the Marmara Sea region of western Turkey reached central Europe via a Balkan route and the Iberian Peninsula via a Mediterranean route. These colonists brought sedentary life, agriculture, and domestic animals and plants to Europe. During their expansion they will have met hunter-gatherers who lived in Europe since the Ice Age, but the two groups mixed initially only to a very limited extent. "They exchanged cultural heritage and knowledge, but rarely spouses," commented anthropologist Joachim Burger, who lead the research. "Only after centuries did the number of partnerships increase."

link.

Terrestrial Effects Of Nearby Supernovae In The Early Pleistocene

Terrestrial Effects Of Nearby Supernovae In The Early Pleistocene

Authors:

Thomas et al

Abstract:

Recent results have strongly confirmed that multiple supernovae happened at distances ~100 pc consisting of two main events: one at 1.7 to 3.2 million years ago, and the other at 6.5 to 8.7 million years ago. These events are said to be responsible for excavating the Local Bubble in the interstellar medium and depositing 60Fe on Earth and the Moon. Other events are indicated by effects in the local cosmic ray (CR) spectrum. Given this updated and refined picture, we ask whether such supernovae are expected to have had substantial effects on the terrestrial atmosphere and biota. In a first cut at the most probable cases, combining photon and cosmic ray effects, we find that a supernova at 100 pc can have only a small effect on terrestrial organisms from visible light, but tropospheric ionization due to the penetration of ≥ TeV cosmic rays will increase by nearly an order of magnitude for thousands of years, and irradiation by muons on the ground and in the upper ocean will increase 20-fold, which will approximately triple the overall radiation load on terrestrial organisms. These effects make possible changes in climate and increased cancer and mutation rates. Further research on the effects of these changes is needed.

How the Mississippi River Drained During the Late Pleistocene

The ancestral Mississippi drainage archived in the late Wisconsin Mississippi deep-sea fan

Authors:

Fildani et al

Abstract:

The response of continental-scale drainage systems to short-term (i.e., millennial-scale) climate change is unknown but has wide implications for understanding climate feedbacks and terrestrial-marine fluxes. The late Wisconsin Mississippi River to deep-sea fan of North America was one of Earth's largest sediment-routing networks during the most recent glacio-eustatic cycle. To understand late Pleistocene sediment production and dispersal related to the partly glaciated, ancestral Mississippi system, we sampled late Wisconsin deep-sea fan channel-fill and lobe deposits for detrital zircon U-Pb and (U-Th)/He double-dating analyses, from Deep Sea Drilling Project (Leg 96) cores and U.S. Geological Survey piston cores. Our results suggest a late Pleistocene glacial Mississippi system that forced a larger transfer of sediment from Cordilleran magmatic provinces and the Canadian Shield when compared to the modern drainage. This indicates a potentially more expansive and/or erosive ancestral Mississippi catchment, and the efficient dispersal of terrigenous sediment, nutrients, and solutes into the deep-sea via high-discharge meltwater and glacial-lake outbursts during ice retreat.

North and South American Pleistocene Quaternary Bears Evolved Giantism Independently

The work of University of Adelaide researchers is shedding new light on the evolution of what are believed to be the largest bears that ever walked the Earth.

Dr Kieren Mitchell, from the University's Australian Centre for Ancient DNA, School of Biological Sciences, has a new paper just published in Biology Letters.

He writes: "During the Pleistocene (2.5 million years ago - 11 thousand years ago) giant bears weighing over 1,000kg roamed both North and South America. These giants belonged to a now practically extinct subfamily of bears - Tremarctinae - which is today only represented by the small, herbivorous Andean spectacled bear (Tremarctos ornatus).

"The giant North American bears (Arctodus) and South American bears (Arctotherium) have long been believed to be each others' closest relative. However, by comparing ancient DNA data obtained from representatives of the two extinct genera and comparing them with the living species, we were able to show that Arctotherium is most closely related to Tremarctos and not Arctodus.

"The implication of this result is that these bears represent a remarkable instance of convergent evolution, as giant bears appear to have evolved independently in both North and South America.

link.

Quaternary glaciation and the Great American Biotic Interchange

Quaternary glaciation and the Great American Biotic Interchange

Authors:

Bacon et al

Abstract:

Recent geological studies demonstrate that the Isthmus of Panama emerged some 10 m.y. earlier than previously assumed. Although absent today in Panama, Central American savanna environments likely developed in connection with the onset of Northern Hemisphere glaciations. As is widely recognized, most of the mammals crossing the isthmus since 2.5 Ma lived in savannas. Could climate-induced vegetational changes across Panama explain the delayed migration of mammals, rather than terrestrial connectivity? We investigate the congruence between cross-continental mammal migration and climate change through analysis of fossil data and molecular phylogenies. Evidence from fossil findings shows that the vast majority of mammals crossed between South and North America after ca. 3 Ma. By contrast, dated mammal phylogenies suggest that migration events started somewhat earlier, ca. 4–3 Ma, but allowing for biases toward greater ages of molecular than geologic dating and uncertainties in the former, we consider this age range not to be significantly earlier than 3 Ma. We conclude that savanna-like environments developed in response to the vast Laurentide ice sheet at the first Quaternary glaciation triggered the initiation of the Great American Biotic Interchange in mammals.

Desert Mammals Disappeared at the Dawn of the Holocene Quaternary From Ethiopia

Paleoenvironmental and biogeographic implications of terminal Pleistocene large mammals from the Ziway–Shala Basin, Main Ethiopian Rift, Ethiopia

Authors:

Lesur et al

Abstract:

The terminal Pleistocene is a period of profound climatic and environmental change in Africa, with paleoenvironmental records documenting tremendous regional variation. Environmental records needed to understand these dynamics are currently lacking from the Horn of Africa. Here we provide a new paleoenvironmental record based of new faunal data from archeological sites along the Bulbula River (Ziway–Shala Basin, Main Ethiopian Rift) in Ethiopia. Research conducted in this area has uncovered archeological assemblages that document various phases of the Late Pleistocene and Early Holocene. Two sites dated to the terminal Pleistocene are considered here, B1s1 and B1s4. The former includes a very large assemblage of more than 20,000 skeletal remains. Both faunal assemblages are dominated by bovids of the tribes Alcelaphini and Antilopini and include Damaliscus hypsodon, a small extinct alcelaphine. The faunal remains from these sites, like those from the contemporary site of Lukenya Hill in south-central Kenya, are indicative of widespread arid grasslands. Several ungulates are found outside of their historic ranges, including Grant's gazelle (Nanger granti) and Dorcas gazelle (Gazella dorcas). Their absence from the region today likely reflects a combination of paleoenvironmental change since the end of the Pleistocene coupled with more recent human impacts. The combination of archeological and faunal data suggests that B1s1 could have been a residential site with seasonal occupation to acquire raw material from local obsidian sources and to hunt bovids in the arid plain, including species that ultimately disappear or shift their range at the onset of the Holocene with the return of warmer and moister conditions.

Smilodon (Sabre Toothed Tiger) Lived on the Plains in South America During the Pleistocene Quaternary

Paleobiology of sabretooth cat Smilodon populator in the Pampean Region (Buenos Aires Province, Argentina) around the Last Glacial Maximum: Insights from carbon and nitrogen stable isotopes in bone collagen

Authors:

Bocherens et al

Abstract:

The sabretooth cat Smilodon populator was the largest felid in South America. It appears in the fossil record in the Early Pleistocene, as an immigrant from North America, and becomes extinct around the Pleistocene–Holocene boundary. The carbon and nitrogen stable isotopic values of collagen were measured for skeletal remains from Smilodon specimens ranging in age from 25 to 10 kyr BP, for the first time in the Pampas region of Argentina. By comparison with similar values obtained on co-eval predators such as Protocyon (large canid) and Panthera onca (jaguar) and a range of potential prey, such as giant ground sloths, glyptodontids, Macrauchenia, Toxodon, equids, cervids, and rodents, it could be established that Smilodon consumed essentially large prey from open landscape, such as Macrauchenia and giant ground sloths during the last 15,000 years of the Late Pleistocene in the Pampa region. It was possibly competing with the large canid Protocyon but the jaguar was apparently feeding on smaller size prey. A more humid climate at the beginning of the Holocene might have been unfavorable to this large predator and could have contributed to its extinction. These results also provide an important insight to understand the ecological processes involved in the Great American Biotic Interchange.

Greatest Megafauna Mammal Extinctions in Europe Began 3,000 Years Ago

Millennial-scale faunal record reveals differential resilience of European large mammals to human impacts across the Holocene

Authors:

Crees et al

Abstract:

The use of short-term indicators for understanding patterns and processes of biodiversity loss can mask longer-term faunal responses to human pressures. We use an extensive database of approximately 18 700 mammalian zooarchaeological records for the last 11 700 years across Europe to reconstruct spatio-temporal dynamics of Holocene range change for 15 large-bodied mammal species. European mammals experienced protracted, non-congruent range losses, with significant declines starting in some species approximately 3000 years ago and continuing to the present, and with the timing, duration and magnitude of declines varying individually between species. Some European mammals became globally extinct during the Holocene, whereas others experienced limited or no significant range change. These findings demonstrate the relatively early onset of prehistoric human impacts on postglacial biodiversity, and mirror species-specific patterns of mammalian extinction during the Late Pleistocene. Herbivores experienced significantly greater declines than carnivores, revealing an important historical extinction filter that informs our understanding of relative resilience and vulnerability to human pressures for different taxa. We highlight the importance of large-scale, long-term datasets for understanding complex protracted extinction processes, although the dynamic pattern of progressive faunal depletion of European mammal assemblages across the Holocene challenges easy identification of ‘static’ past baselines to inform current-day environmental management and restoration.

Paleolithic Papers #3

Modern Humans (Homo sapiens):

A modified bear bone from 12,500 years ago near the Pleistocene/Holocene boundary pushes back the time of the earliest occupants of Ireland.

There is some contention over the Pleistocene reports of the earliest domestic dogs. A flag raised has been there has been no study of wolf variation at the time and those early dogs might just be wolves rather than domesticated dogs.

DNA sequencing of a modern human from Ust'-Ishim gives some interesting insights.

The Water Canyon site from Pleistocene/Holocene Boundary New Mexico provides some interesting insights to the PaleoIndians of North America.

The latest in a series of transformative studies of DNA from prehistoric Europeans focuses on mitochondrial DNA, bringing fresh surprises and filling in important details of the early stages of a European ancestry stretching back more than 40,000 years.

John Hawks grumps over the comments about the exact number of times archaic human populations and modern humans interbred.

Lezetxiki Hominin (Homo neanderthalensis?):

The humerus of the Lezetxiki Hominin shows affinities with the Pit of Bones hominins (that appears to be early Neandertals and/or Denisovans). If so, this may represent an early Neandertal, too.  It dates from 164 thousand years ago (+/- 9k years).

Denisovans (no species name assigned yet):

Denisovans, Neandertals and modern humans shared an ancestor about a million years ago.  Not counting the introgressions since then.

Hominins (general): 

When and how did hominins become bipedal and lose their hair?

The Siberian Unicorn Found a Refugium, Surviving until 29,000 Years ago

The beautiful title "Siberian unicorn" belongs to Elasmotherium sibiricum - an elasmotherium Siberian rhinoceros, which as previously thought became extinct 350,000 years ago. Nowadays the researchers of Tomsk State University (TSU) figured out that the "unicorn" found his last refuge "only" 29,000 years ago in Kazakhstan. The article, describing the new location of the fossil mammals in the Pavlodar Irtysh, was published in February 2016 in the American Journal of Applied Science.

"Most likely, in the south of Western Siberia it was a refúgium, where this rhino had preserved the longest in comparison with the rest of its range. There is another option that it could migrate and dwell for a while on the more southern areas," said Andrey Shpanski, a paleontologist at TSU. These conclusions were made due to research of the rhinocero's skull, found near Kozhamzhar village in Pavlodar region (Kazakhstan). The skull is well preserved: there are some cracks but no trace of pelletization, gnawing, and exfoliation. The fossils of the "unicorn" were examined by radiocarbon AMS-method analysis in the laboratory 14CHRONO Centre for Climate, the Environment, and Chronology (School of Geography, Archaeology and Palaeoecology; Queen's University Belfast; Belfast, UK). It turned out that the skull belonged to the animals that died 29,000 years ago. "Most likely, it was a very large male of very large individual age (teeth not preserved). The dimensions of this rhino today are the biggest of those described in the literature, and the proportion are typical," said the University's scientist.

link.

I Smell a Coming Academic Bun Fight: Sima de los Huesos Hominins Really Were Early Neandertals

Previous analyses of the hominins from Sima de los Huesos in 2013 showed that their maternally inherited mitochondrial DNA was distantly related to Denisovans, extinct relatives of Neandertals in Asia. This was unexpected since their skeletal remains carry Neandertal-derived features. Researchers of the Max Planck Institute for Evolutionary Anthropology in Leipzig, Germany, have since worked on sequencing nuclear DNA from fossils from the cave, a challenging task as the extremely old DNA is degraded to very short fragments. The results now show that the Sima de los Huesos hominins were indeed early Neandertals. Neandertals may have acquired different mitochondrial genomes later, perhaps as the result of gene flow from Africa.

Until now it has been unclear how the 28 400,000-year-old individuals found at the Sima de los Huesos ("pit of bones") site in Northern Spain were related to Neandertals and Denisovans who lived until about 40,000 years ago. A previous report based on analyses of mitochondrial DNA from one of the specimens suggested a distant relationship to Denisovans, which is in contrast to other archaeological evidence, including morphological features that the Sima de los Huesos hominins shared with Neandertals.

link.

The Importance of Atmospheric Carbon in the end of the Last Glaciation

As the Earth emerged from its last ice age several thousand years ago, atmospheric carbon dioxide increased and further warmed the planet. Scientists have long speculated that the primary source of this CO2 was from the deep ocean around Antarctica, though it has been difficult to prove.

A new study published this week in Proceedings of the National Academy of Sciences confirmed that the ocean played a significant role in the rise of atmospheric carbon dioxide, but also documents the signature of land-based carbon sources in Antarctic ice cores that contributed to abrupt increases in CO2.

"There wasn't a steady rate of rising carbon dioxide during the last deglaciation," said Edward Brook, an Oregon State University paleoclimatologist and co-author on the PNAS study. "It happened in fits and starts. With the new precise techniques we developed to fingerprint the sources, it is apparent that the early carbon largely came from the ocean, but we think the system got a jolt from an influx of land-based carbon a few times as the climate warmed."

link.

Yes, Neandertals Were Omnivores

Ecological niche of Neanderthals from Spy Cave revealed by nitrogen isotopes of individual amino acids in collagen

Authors:

Naito et al

Abstract:

This study provides a refined view on the diet and ecological niche of Neanderthals. The traditional view is that Neanderthals obtained most of their dietary protein from terrestrial animals, especially from large herbivores that roamed the open landscapes. Evidence based on the conventional carbon and nitrogen isotopic composition of bulk collagen has supported this view, although recent findings based on plant remains in the tooth calculus, microwear analyses, and small game and marine animal remains from archaeological sites have raised some questions regarding this assumption. However, the lack of a protein source other than meat in the Neanderthal diet may be due to methodological difficulties in defining the isotopic composition of plants. Based on the nitrogen isotopic composition of glutamic acid and phenylalanine in collagen for Neanderthals from Spy Cave (Belgium), we show that i) there was an inter-individual dietary heterogeneity even within one archaeological site that has not been evident in bulk collagen isotopic compositions, ii) they occupied an ecological niche different from those of hyenas, and iii) they could rely on plants for up to ∼20% of their protein source. These results are consistent with the evidence found of plant consumption by the Spy Neanderthals, suggesting a broader subsistence strategy than previously considered.

Homo habilis & Paranthropus boisei Pleistocene Quaternary Habitat was NOT the Garden of Eden

Scientists have pieced together an early human habitat for the first time, and life was no picnic 1.8 million years ago.

Our human ancestors, who looked like a cross between apes and modern humans, had access to food, water and shady shelter at a site in Olduvai Gorge, Tanzania. They even had lots of stone tools with sharp edges, said Gail M. Ashley, a professor in the Rutgers Department of Earth and Planetary Sciences in the School of Arts and Sciences.

But "it was tough living," she said. "It was a very stressful life because they were in continual competition with carnivores for their food."

During years of work, Ashley and other researchers carefully reconstructed an early human landscape on a fine scale, using plant and other evidence collected at the sprawling site. Their pioneering work was published recently in the Proceedings of the National Academy of Sciences.

The landscape reconstruction will help paleoanthropologists develop ideas and models on what early humans were like, how they lived, how they got their food (especially protein), what they ate and drank and their behavior, Ashley said.

link.

Was There a Significant Human Influence on the Flora of North Africa During the Pleistocene & Holocene?

Pleistocene and Holocene herbivore diets and palaeoenvironments in the Gebel Akhdar (Libya): Implications for past human populations

Authors:

Reade et al

Abstract:

The Gebel Akhdar massif in Cyrenaica, northeast Libya, has yielded a long record of human occupation going back at least 100,000 years. To date, there is only a limited understanding of how the landscape of the region varied in response to the climatic fluctuations of the last glacial–interglacial cycle, and the implications of these changes for local human populations remain largely unexplored. This study provides an isotope-based interpretation of past environments directly linked to the archaeological record. Tooth enamel stable carbon isotope ratios (δ13C) from herbivore species hunted by past human populations are used to infer the isotopic characteristics of past diet and vegetation, and in turn the likely environmental conditions that prevailed during periods when humans were active within the landscape. To provide a baseline from which to interpret the archaeological δ13C data, modern samples are considered in relation to their diet and environmental origin. Archaeological samples come from 2 cave sites, Haua Fteah and Hagfet ed Dabba, and span a period from oxygen isotope stage 4 to the mid-Holocene. Whilst results indicate a more arid environment in the Pleistocene and an increase in humidity at the onset of the Holocene, the overall picture is one of relative environmental stability. The biggest landscape change observed in the data occurs during the mid-Holocene Neolithic, when C4 plant species become evident in the herbivore diet for the first time. There is little evidence to suggest that this occurred at a time of any large-scale climate variation, and thus the contribution of anthropogenic influences to vegetation change is considered likely.