Ecological Changes During the Glacial & Interglacials of the late Paleozoic ice age of Pennsylvanian Carboniferous Argentina

Compositional turnover and ecological changes related to the waxing and waning of glaciers during the late Paleozoic ice age in ice-proximal regions (Pennsylvanian, western Argentina)

Author:

Balseiro

Abstract:

The late Paleozoic ice age (LPIA) had a profound effect on the biota. Despite much research having been focused on paleotropical regions or global-scale analyses, regional ecological changes have seldom been studied in ice-proximal basins. Here, I study the compositional turnover and diversity structure across the main Carboniferous glacial event recorded in western Argentina and the subsequent nonglacial interval. Brachiopod and bivalve data from western Argentina suggest that the transition from glacial to nonglacial climates caused major compositional changes. Turnover, however, was not uniform across the bathymetric gradient, being higher in deep environments. Because extirpation was concentrated in brachiopods, but immigration was similar in both clades, the taxonomic structure of the region was significantly modified. Although regional hierarchical diversity structure and occupancy distributions remained stable, dissecting the analysis in brachiopods and bivalves underscores that both clades had different responses to climate change. Brachiopods, on the one hand, show stability in the diversity structure and a very slight decrease in occupancies of intermediate genera, while bivalves show an important rise in diversity, both at the environment and regional scale, and an increase in genera with intermediate occupancies. The bathymetric diversity gradient was also modified from hump shaped with maximum diversity in the deep subtidal to a linear gradient with maximum values toward the offshore. However, relative compositional differences within environments remained stable, with maximum values at intermediate depths both in glacial and nonglacial intervals. Moreover, local-scale coexistence between brachiopods and bivalves changed in the nonglacial interval, showing significant segregation, which indicates relevant modifications in community assembly dynamics. Results from western Argentina highlight the magnitude of regional-scale ecological changes during the LPIA in ice-proximal regions, suggesting that the waxing and waning of glaciers was able to cause regional taxonomic turnover and medium-scale ecological changes even during intervals of relative macroevolutionary quiescence.

The Bashkirian Pennsylvanian Carboniferous was the Peak of the Late Paleozoic Ice Age

Ice volume and paleoclimate history of the late Paleozoic ice age from conodont apatite oxygen isotopes from naqing (Guizhou, China)

Authors:

Chen et al

Abstract:

A high-resolution and continuous conodont apatite oxygen isotope record spanning the late Viséan to Middle Permian is reported from South China, which is interpreted with respect to the ice volume and/or tropical sea water temperature history of the Late Paleozoic Ice Age (LPIA). The presented δ18O record shows significant fluctuations in δ18O from the late Viséan to Middle Permian with highest values observed in the Bashkirian (Early Pennsylvanian). The δ18O maximum coincides with a major eustatic sea level fall recorded in low-latitudinal successions, and postdates the significant increases in 87Sr/86Sr and δ13Ccarb measured on well-preserved brachiopod calcite, which can be interpreted as reflecting intensified weathering as consequence of the closure of Rheic Ocean as well as enhanced carbon burial. Both processes may have contributed to lower greenhouse gas levels and cooled down the Earth's surface, triggering the maximum glaciation. The high Bashkirian δ18O values are interpreted to represent the glacial maximum of the LPIA. A coeval change in faunal composition and a decreasing diversity in climate-sensitive marine invertebrates can be ascribed to icehouse cooling and/or loss of habitat. Despite inconsistenceies with earlier interpretations that the Early Permian represented the glacial maximum of the LPIA as inferred from Gondwanan glacial sediments records, the suggested Bashkirian glacial maximum agrees well with ice extent estimates based on the regional tectonic history in Gondwana, which suggests that the Bashkirian glaciation occurred during Gondwana interior uplift promoting maximum ice cover of the entire LPIA. However, maximum glaciation is only poorly represented in the depositional record because large parts of the glacial deposits were possibly removed by erosion as outlined by a major regional unconformity.

A Marine Incursion in Pennsylvanian Carboniferous Canada and its Impacts

A Marine Incursion In the Lower Pennsylvanian Tynemouth Creek Formation, Canada: Implications For Paleogeography, Stratigraphy and Paleoecology

Authors:

Falcon-Lang et al

Abstract:

We document the occurrence of a marine bed, and its associated biota, in the Lower Pennsylvanian (Langsettian) Tynemouth Creek Formation of New Brunswick, and discuss its implications for paleogeography, stratigraphy, and paleoecology. This is only the second marine interval found in the entire Pennsylvanian fill of the Maritimes Basin of Canada, the other being recently found in the broadly same-age Joggins Formation of Nova Scotia. Evidence for the new marine transgression comprises an echinoderm-rich limestone that infills irregularities on a vertic paleosol surface within the distal facies of a syntectonic fluvial megafan formed under a seasonally dry tropical climate. Gray, platy ostracod-rich shales and wave-rippled sandstone beds that directly overlie the marine limestone contain trace fossils characteristic of the Mermia Ichnofacies, upright woody trees, and adpressed megafloras. This association represents bay-fills fringed by freshwater coastal forests dominated by pteridosperms, cordaites, and other enigmatic plants traditionally attributed to dryland/upland habitats. The fossil site demonstrates that marine transgressions extended farther into the interior of Pangea than has previously been documented, and may allow correlation of the Tynemouth Creek and Joggins Formations with broadly coeval European successions near the level of the Gastrioceras subcrenatum and G. listeri marine bands. It also helps explain the close similarity of faunas between the Maritimes Basin and other paleotropical basins, if transgressions facilitated migration of marine taxa into the continental interior.

The Climate of Moscovian Carboniferous Czech Republic

Middle Moscovian climate of eastern equatorial Pangea recorded in paleosols and fluvial architecture

Authors:

Opluštil et al

Abstract:

Fluvial architecture in conjunction with the morphology, petrographical, mineralogical and geochemical composition of intercalated paleosols of the Middle Moscovian upper Radnice Member of the continental Kladno-Rakovník Basin, Czech Republic, were studied in order to decipher climatic signals in the eastern Pangea region during the Late Paleozoic Ice Age. Analysis of architectural elements reveals vertical changes from strongly amalgamated channel fills of bedload-dominated braided streams, to isolated ribbon-like meandering channel fills in mud-dominated floodplain strata. The resulting large scale fining upward cycles separated by laterally widespread bounding surfaces are interpreted as having been formed in response to allocyclic processes driven by variable intensities of precipitation which affected the density of vegetation cover across the landscape and, in turn, the amount and character of sediment supply. Bedload-dominated parts of the cycles were deposited during periods of low water tables and sparse vegetation, whereas periods of maximum precipitation resulted in higher water tables and a greater density of vegetation cover, which, in turn stabilized the floodplains and fixed streams to narrow meandering/anastomosing channels in mud-dominated floodplain strata. Associated paleosols resemble modern Vertisols, which form under seasonal precipitation in modern settings. However, coeval formation of inertinite-rich peat represented by histosols and gleyed low chroma Vertisols in the stratigraphically lowest paleosol succession occurred under overlapping marginal conditions for the climatically different types of soils (i.e., under a seasonal climate where precipitation exceeds evapotranspiration for 6 to 8 months out of the year). Mean annual precipitation calculated from the geochemistry of B horizons of Vertisols (CALMAG proxy) varies between 1815 and 1730 mm/yr.

The Role of Continental Drift & Glaciations on Carbonifierous PaleoClimates

Impact of continental motion and dynamic glaciations on low-latitude climate during the Carboniferous: The record of the Wyoming Shelf (Western United States)

Authors:

Blanchard et al

Abstract:

The dynamic character of the Late Paleozoic Ice Age is evident from glacial deposits, but its impact on tropical climate is not well constrained. Global changes in climate are overprinted on longer-term paleogeographic variations, resulting in a complex time–space distribution of climate-sensitive lithologies. The significance of such lithologies in Carboniferous successions of the western United States has not been fully explored. In this study, we provide new interpretations for the paleoclimatic context of the Amsden and Tensleep Formations (Pennsylvanian, Northern Wyoming, USA). The Amsden Formation consists of a basal sandstone member overlain by red siltstones containing pisolites. Very large-scale (~ 10 m) cross-bedding within the basal sandstone indicates deposition in an erg environment. Iron pisoid-rich layers in the overlying member suggest an evolution toward more humid conditions. Persistent arid conditions during the middle Pennsylvanian are suggested by eolian sandstones and calcretes in the overlying Tensleep Formation. These formations were deposited on the karst topography that developed on top of the lower to middle Mississippian Madison Group. Although the development of karstic features implies that humid conditions prevailed during the late Mississippian, evaporites and evidence for early dolomitization within the formation suggest that it was deposited under arid conditions. These relationships argue for a long-term climate evolution from arid to humid during the Mississippian, and a return to arid conditions during the Pennsylvanian. This trend can be explained by the northward drift from ~ 15°S to ~ 12°N. A comparison with contemporaneous records reveals a diachronous evolution across western Pangaea, with the climatic conditions documented on the Wyoming Shelf being reached later in eastern North America. These relationships indicate that plate motion considerably overprints long-term climatic records. Departures from this trend, suggested by the presence of erg deposits in the basal Amsden Formation, record the overprinting of shorter periods of climate change.

The Flora of Asselian Permian Equitorial and Seasonally dry Pangea...From Las Cruces, NM

Early Permian (Asselian) vegetation from a seasonally dry coast in western equatorial Pangea: Paleoecology and evolutionary significance

Authors:

Falcon-Lang et al

Abstract:

The Pennsylvanian–Permian transition has been inferred to be a time of significant glaciation in the Southern Hemisphere, the effects of which were manifested throughout the world. In the equatorial regions of Pangea, the response of terrestrial ecosystems was highly variable geographically, reflecting the interactions of polar ice and geographic patterns on atmospheric circulation. In general, however, there was a drying trend throughout most of the western and central equatorial belt. In western Pangea, the climate proved to be considerably more seasonally dry and with much lower mean annual rainfall than in areas in the more central and easterly portions of the supercontinent. Here we describe lower Permian (upper Asselian) fossil plant assemblages from the Community Pit Formation in Prehistoric Trackways National Monument near Las Cruces, south-central New Mexico, U.S.A. The fossils occur in sediments within a 140-m-wide channel that was incised into indurated marine carbonates. The channel filling can be divided into three phases. A basal channel, limestone conglomerate facies contains allochthonous trunks of walchian conifers. A middle channel fill is composed of micritic limestone beds containing a brackish-to-marine fauna with carbon, oxygen and strontium isotopic composition that provide independent support for salinity inferences. The middle limestone also contains a (par)autochthonous adpressed megaflora co-dominated by voltzian conifers and the callipterid Lodevia oxydata. The upper portions of the channel are filled with muddy, gypsiferous limestone that lacks plant fossils. This is the geologically oldest occurrence of voltzian conifers. It also is the westernmost occurrence of L. oxydata, a rare callipterid known only from the Pennsylvanian–Permian transition in Poland, the Appalachian Basin and New Mexico. The presence of in situ fine roots within these channel-fill limestone beds and the taphonomic constraints on the incorporation of aerial plant remains into a lime mudstone indicate that the channel sediments were periodically colonized by plants, which suggests that these species were tolerant of salinity, making these plants one of, if not the earliest unambiguous mangroves.

Evidence of Arthropod Herbivory and Detrivory From Bolsovian/Asturian Pennsylvanian Carboniferous Britain

HERBIVOROUS AND DETRITIVOROUS ARTHROPOD TRACE FOSSILS ASSOCIATED WITH SUBHUMID VEGETATION IN THE MIDDLE PENNSYLVANIAN OF SOUTHERN BRITAIN

Authors:

Falcon-Lang et al

Abstract:

We describe plant–arthropod associations from the Middle Pennsylvanian (late Bolsovian–early Asturian) Pennant Sandstone Formation of southern Britain. Our material comprises calcified cordaitaleans and tree-fern axes, preserved in braided channel deposits, and interpreted as remains of subhumid riparian vegetation distinct from that of coeval coal swamps. The first plant–arthropod association, attributed to herbivorous insects, comprises cambial damage to cordaitalean leafy branches, resulting in traumatic wound response. The second and most widespread association, attributable to detritivorous oribatid mites, includes tunnels and galleries containing widely scattered, clustered, or densely packed microcoprolites within the inner root mantle of marattialean tree ferns and cordaitalean trunks and branches. Diameter data for tunnels and microcoprolites are multimodal, recording four or five instars of oribatid mites that parallel instar-based fecal pellet and body lengths in modern taxa. The third association attributed, possibly, to an arthropleurid, comprises a single, very large (19 × 14 mm) coprolite. Included plant fragments support a previous conjecture that arborescent lycopsids formed part of this iconic arthropod's diet. Mucus-lined burrows within the macrocoprolite imply that fecal material was processed by annelids. The high diversity and frequency of plant–arthropod associations are unusual for Mid-Pennsylvanian time, and may reflect previously undetected interactions in those ecosystems that lay outside “coal forest swamps.”

Color Vision in the Paleozoic: Fossil Cones and Rods Found in Gzhelian Pennsylvannian Carboniferous Fish

The finding reveals that such eye cells have existed for at least 300 million years, and that the ancient fish they were discovered in likely saw in color, according to the study's scientists.

Human vision depends on pigments that absorb light. These pigments lie inside cells known as rods and cones. Cones are sensitive to color and also help perceive fine detail and rapid changes. Rods are more sensitive to light than cones, but are not sensitive to color, and are responsible for peripheral and night vision. Both rods and cones are found in a layer of tissue in the back of the eye known as the retina. [Vision Quiz: What Can Animals See?]

Myllokunmingia may be one of the earliest known creatures with a backbone, and this creature may have possessed a rudimentary cameralike eye, which suggests vision dates back at least 520 million years. However, much remains unknown about the evolution of vision, since the soft tissue of the eye usually decays rapidly after death.

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Erpetonyx arsenaultorum: a new and Oldest Known Parareptile From Gzhelian Pennsylvannian Carboniferous Prince Edward Island

The oldest parareptile and the early diversification of reptiles

Authors:

Modesto et al

Abstract:

Amniotes, tetrapods that evolved the cleidoic egg and thus independence from aquatic larval stages, appeared ca 314 Ma during the Coal Age. The rapid diversification of amniotes and other tetrapods over the course of the Late Carboniferous period was recently attributed to the fragmentation of coal-swamp rainforests ca 307 Ma. However, the amniote fossil record during the Carboniferous is relatively sparse, with ca 33% of the diversity represented by single specimens for each species. We describe here a new species of reptilian amniote that was collected from uppermost Carboniferous rocks of Prince Edward Island, Canada. Erpetonyx arsenaultorum gen. et sp. nov. is a new parareptile distinguished by 29 presacral vertebrae and autapomorphies of the carpus. Phylogenetic analyses of parareptiles reveal E. arsenaultorum as the closest relative of bolosaurids. Stratigraphic calibration of our results indicates that parareptiles began their evolutionary radiation before the close of the Carboniferous Period, and that the diversity of end-Carboniferous reptiles is 80% greater than suggested by previous work. Latest Carboniferous reptiles were still half as diverse as synapsid amniotes, a disparity that may be attributable to preservational biases, to collecting biases, to the origin of herbivory in tetrapods or any combination of these factors.

Deserts in Southwest Gondwana During Moscovian Pennsylvannian Carboniferous

Late Pennsylvanian aridification on the southwestern margin of Gondwana (Paganzo Basin, NW Argentina): A regional expression of a global climate perturbation
Authors:

Gulbranson et al

Abstract:

Paleosols of the Paganzo Group preserve a shift from the cool climate of the glaciated high-latitudes of Gondwana to a long-term aridification punctuated by shorter-term fluctuations in moisture regime in the late Carboniferous. A quantitative reconstruction of paleoclimate inferred from the major element chemistry of paleosol horizons and the stable isotope geochemistry of soil-formed minerals and organic matter indicates that the climate varied substantially during deglaciation along the southwestern margin of Gondwana in the Middle Pennsylvanian including two geologically abrupt shifts to notably drier conditions. This dynamic climate terminates in a dry subhumid moisture regime characterized by steppe/dry scrub floral provinces. Quantitative estimates of net primary production during the post-glacial interval inferred from paleosol major element geochemistry are consistent with a mesic soil temperature regime, in agreement with paleotemperatures inferred from pedogenic goethite δD and δ18O. Contemporaneous aridification is well expressed in paleotropical continental records with Middle to Late Pennsylvanian aridification coincident with an abrupt evolutionary change in many marine and terrestrial species. The synchrony of aridification between the high and low paleo-latitudes and concomitant major restructuring of some ecosystems suggests that a paleoclimate change of global nature occurred during the Moscovian and Kasimovian.

Westphalian Pennsylvanian Carboniferous Trackways From Morocco

First Occurrence of Tetrapod Footprints from Westphalian Strata of the Sidi Kassem Basin, Central Morocco

Authors:

Lagnaoui et al

Abstract:

The Sidi Kassem Basin is the only limnic basin of Westphalian age in Morocco. It is built up of 1,250 m of alluvio-fluvial to lacustrine deposits that have so far yielded plant fossils and invertebrate remains only. Recent exploration for fossil tetrapod footprints in floodplain-deposits of the basin revealed a moderately diverse vertebrate ichnofauna composed of tracks assigned to cf. Batrachichnus Woodworth, 1900; cf. Hylopus Dawson, 1882; Dimetropus Romer and Price, 1940; and Notalacerta Butts, 1891. The tracks can be referred to temnospondyl, anamniote reptiliomorph, non-therapsid synapsid (pelycosaur), and captorhinomorph trackmakers. The described ichnoassemblage is important in at least three aspects: (1) It suggests an Early to mid-Pennsylvanian age for the footprint-bearing strata of the study area. (2) It is the oldest association of tetrapod footprints from Africa. (3) It is the first evidence of the relatively rare ichnogenera cf. Hylopus and Notalacerta outside of North America and Europe. Judged from the variety of tetrapod tracks and previously collected floral remains, the Sidi Kassem Basin must have represented a well-established continental ecosystem during Pennsylvanian time. Further exploration for trace and body fossils of Palaeozoic vertebrates in this basin may be important for the reconstruction of early tetrapod evolution.

Pennsylvanian Carboniferous Synapsid Ianthodon schultzei Revisited

New information on the cranial and postcranial anatomy of the early synapsid Ianthodon schultzei (Sphenacomorpha: Sphenacodontia), and its evolutionary significance

Authors:

Spindler et al

Abstract:

Newly identified material belonging to the holotype specimen of Ianthodon schultzei substantially increases our knowledge of this poorly known basal sphenacodont synapsid from the fossil site in Garnett, Kansas (Missourian, Late Pennsylvanian). The original description, based on a partial dermal skull roof, is augmented with information on the palate and braincase, together with data on the mandible and a few postcranial elements. The known skeletal morphology resembles that of Haptodus garnettensis, another synapsid taxon known from this locality, but with fewer marginal, distinctly recurved teeth and smaller teeth on the transverse flange of the pterygoid. Although recognizing that the holotype and only known specimen represents a juvenile individual, Ianthodon appears to reflect a more basal sphenacodontian condition than H. garnettensis. A restricted phylogenetic analysis based on previous work and newly scored characters for Ianthodon, Cutleria and Pantelosaurus supports this hypothesis. The Garnett locality appears to preserve an assemblage of synapsids (Haptodus, Ianthasaurus, Ianthodon) that are close to the base of the large clade that includes Edaphosauridae and Sphenacodontia, suggesting that an initial diversification of this clade occurred well within the Carboniferous Period.

Just What WAS the "Parrsboro jaw" From Pennsylvanian Carboniferous Nova Scotia

Redescription and Phylogenetic Analysis of the Mandible of an Enigmatic Pennsylvanian (Late Carboniferous) Tetrapod from Nova Scotia, and the Lability of Meckelian Jaw Ossification

Authors:

Sookias et al

Abstract:

The lower jaw of an unidentified Pennsylvanian (Late Carboniferous) tetrapod from Nova Scotia – the “Parrsboro jaw”- is redescribed in the light of recent tetrapod discoveries and work on evolution of tetrapod mandibular morphology and placed for the first time in a numerical cladistics analysis. All phylogenetic analyses place the jaw in a crownward polytomy of baphetids, temnospondyls, and embolomeres. Several features resemble baphetids and temnospondyls including dermal ornamentation, absence of coronoid teeth, and presence of coronoid shagreen. Dentary dentition is most similar to Baphetes. An adsymphysial toothplate may not preclude temnospondyl affinity. An apparent large exomeckelian fenestra, with the dorsal foraminal margins formed by an unossified element, echoes the morphology of the stem tetrapod Sigournea and is unusually primitive given the other features of the jaw. The jaw may thus provide an example of an intermediate stage in Meckelian element evolution.

Evidence of Warming and Cooling Events From the Pennsylvanian Carboniferous to the Permian

Warm water benthic foraminifera document the Pennsylvanian–Permian warming and cooling events — The record from the Western Pangea tropical shelves

Author:

Davydov

Abstract:

Shallow warm water benthic foraminifera (SWWBF), including all larger fusulinids (symbiont-bearing benthic foraminifera), are among the best indicators of paleoclimate and paleogeography in the Carboniferous and Permian. The distribution of benthic foraminifera in space and time constrain important tectonic, paleogeographic and climatic events at a global scale. The North American shelves during Pennsylvanian and Permian time – though geographically within the tropical belt – are characterized by temperate environments with significantly lower foraminifera diversification and rare occurrences of warm water Tethyan forms, that are in general appear in the region as a migration entities. Such environments allow documentation of warming episodes associated with sudden immigration of warm water and exotic forms of SWWBF that evolved elsewhere into the area. First occurrence datum (FOD) of the forms exotic to North America during warming episodes are always delayed in respect of their First Appearance Datum (FAD) elsewhere. The time of delay and taxonomic diversity of fusulinids in North America shelves depended on the scale and intensity of the warming episodes. Cooling events, on the other hand, are associated with decreased taxonomic diversity and appearances of endemic forms characteristic only of temperate water provinces. The occurrence of these forms in Boreal and North American provinces appears to be isochronous, as their environments are uniform and induce their uniform and isochronous distribution. Several warming and cooling episodes during Pennsylvanian–Permian time are recognized. The differences between taxonomic variations in each event could potentially be used for provisional estimation of the degree of climatic change. A strong link between biotic and climatic events in North American province and the similarity of biotic changes in the North American and other provinces suggests that paleoclimatic events in North American province were controlled by global factors.

Evidence Temnospodyl Micromelerpeton Regenerated Limbs From Gzhelian Pennsylvanian Carboniferous

Early evolution of limb regeneration in tetrapods: evidence from a 300-million-year-old amphibian

Authors:

Fröbisch et al

Abstract:

Salamanders are the only tetrapods capable of fully regenerating their limbs throughout their entire lives. Much data on the underlying molecular mechanisms of limb regeneration have been gathered in recent years allowing for new comparative studies between salamanders and other tetrapods that lack this unique regenerative potential. By contrast, the evolution of animal regeneration just recently shifted back into focus, despite being highly relevant for research designs aiming to unravel the factors allowing for limb regeneration. We show that the 300-million-year-old temnospondyl amphibian Micromelerpeton, a distant relative of modern amphibians, was already capable of regenerating its limbs. A number of exceptionally well-preserved specimens from fossil deposits show a unique pattern and combination of abnormalities in their limbs that is distinctive of irregular regenerative activity in modern salamanders and does not occur as variants of normal limb development. This demonstrates that the capacity to regenerate limbs is not a derived feature of modern salamanders, but may be an ancient feature of non-amniote tetrapods and possibly even shared by all bony fish. The finding provides a new framework for understanding the evolution of regenerative capacity of paired appendages in vertebrates in the search for conserved versus derived molecular mechanisms of limb regeneration.

Eocasea martini: a new Carnivorous Caseid Synapsid From Pennsylvanian Carboniferous Kansas

The Oldest Caseid Synapsid from the Late Pennsylvanian of Kansas, and the Evolution of Herbivory in Terrestrial Vertebrates

Authors:

Reisz et al

Abstract:

The origin and early evolution of amniotes (fully terrestrial vertebrates) led to major changes in the structure and hierarchy of terrestrial ecosystems. The first appearance of herbivores played a pivotal role in this transformation. After an early bifurcation into Reptilia and Synapsida (including mammals) 315 Ma, synapsids dominated Paleozoic terrestrial vertebrate communities, with the herbivorous caseids representing the largest vertebrates on land. Eocasea martini gen. et sp. nov., a small carnivorous caseid from the Late Carboniferous, extends significantly the fossil record of Caseidae, and permits the first clade-based study of the origin and initial evolution of herbivory in terrestrial tetrapods. Our results demonstrate for the first time that large caseid herbivores evolved from small, non-herbivorous caseids. This pattern is mirrored by three other clades, documenting multiple, independent, but temporally staggered origins of herbivory and increase in body size among early terrestrial tetrapods, leading to patterns consistent with modern terrestrial ecosystem.

Reassessing the Extent of Serpukhovian to Bashkirian Carboniferous Southwestern Gondwana's Glaciation

Reassessment of mid-Carboniferous glacial extent in southwestern Gondwana (Rio Blanco Basin, Argentina) inferred from paleo-mass transport of diamictites

Authors:

Gulbranson et al

Abstract:

Late Paleozoic glacial diamictites occur in many localities in western Argentina, indicating that the region was strongly affected by glaciation during the mid-Carboniferous (late Serpukhovian–early Bashkirian). In most instances these diamictites are found in steeply walled paleovalley settings in the Andean Precordillera. This study presents new data from a locality north of the Precordillera that suggests an additional, distinct, volume of ice existed in the region during the Carboniferous. The glacigenic diamictites in the Rio Blanco Basin were ultimately emplaced as gravity flows, precluding inferences of paleo-ice volume. Fold nose orientation and soft-sediment groove orientations within the diamictites indicate that the deposits were emplaced from north to south, suggesting that glacial ice was most likely not sourced from the proto-Precordillera at this locality, requiring the need for another ice center to the north of the basin. Diamictite facies indicates that the sediment was initially supplied to the study area by a warm-based glacier.

The Late Paleozoic Deglaciation Transition in Southwestern Gondwana

The late Paleozoic El Imperial Formation, western Argentina: Glacial to post-glacial transition and stratigraphic correlations with arc-related basins in southwestern Gondwana

Authors:

Henry et al

Abstract:

The El Imperial Formation of the San Rafael Basin records a succession of depositional environments during the latest Mississippian to earliest Permian that span before, during, and after the glaciation of west central Argentina. At the base of the formation, a restricted marine environment is recorded in mudstone containing marl and rippled and deformed sandstone beds. This unit, or sequence 1, is incised by a deltaic facies association composed of cross-bedded sandstone and conglomerate that form at least 5 stacked Gilbert deltas. The deltaic facies association grades upward into the glacially-influenced facies association, made up of stratified diamictite, mudstone with dropstones, and massive deformed sandstone, indicating deposition by wet-based tidewater glaciers that calved icebergs into the basin, with contributions from mass movement processes. The glacially-influenced facies association is overlain by mudstone and horizontally laminated and cross-bedded sandstone of the post-glacial open marine facies association, recording post-glacial transgression followed by relative sea level fall. The deltaic, glacially-influenced, and post-glacial open marine facies associations comprise sequence 2. Sequence 2 is incised by conglomerate of the upper fluvial member, or sequence 3.

The strata of the El Imperial Formation are correlated to those of the other arc-related basins of western Argentina: Río Blanco, Calingasta–Uspallata, and Tepuel. A Bashkirian transgression and fluvial incision in the El Imperial Formation correlate with events in the Río Blanco and Calingasta–Uspallata Basins to the north, whereas glaciation continues to the south in the Tepuel Basin through the Early Permian. The deviating stratigraphic record of the Tepuel Basin may be the result of its higher latitudinal position during the Pennsylvanian–Early Permian and higher altitude due to either tectonic convergence of the Patagonian microplate or convergence along the Panthalassan margin of southwestern Gondwana.

Like Salmon in Reverse, Carboniferous Freshwater Sharks Returned to the Sea to Breed

Like salmon in reverse, long-snouted Bandringa sharks migrated downstream from freshwater swamps to a tropical coastline to spawn 310 million years ago, leaving behind fossil evidence of one of the earliest known shark nurseries.

That's the surprising conclusion of University of Michigan paleontologist Lauren Sallan and a University of Chicago colleague, who reanalyzed all known specimens of Bandringa, a bottom-feeding predator that lived in an ancient river delta system that spanned what is today the Upper Midwest.

The new findings, scheduled for online publication Jan. 7 in the Journal of Vertebrate Paleontology, mark the earliest known example of shark migration—a behavior that persists today among species such as tiger sharks in Hawaii.

The Bandringa fossils, as reinterpreted by Sallan and Michael Coates, also reveal the only known example of a freshwater to saltwater shark migration, as well as the earliest example of a shark nursery where fossilized egg cases and juvenile sharks were preserved in the same sediments.

"This pushes migratory behavior in sharks way back," said Sallan, an assistant professor in the U-M Department of Ecology and Evolutionary Biology. "These sharks bred in the open ocean and spent the rest of their lives in fresh water. No shark alive today is known to do that."

The long-extinct Bandringa is likely one of the earliest close relatives of modern sharks. It resembled present-day sawfish and paddlefish, with a spoon-billed snout up to half its body length. Juveniles were 4 to 6 inches long and grew into adults of up to 10 feet.

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Climate and Eustasy Driven Cycles in Pennsylvanian Carboniferous Donets Basin, Ukraine

Climate- and eustasy-driven cyclicity in Pennsylvanian fusulinid assemblages, Donets Basin (Ukraine)

Authors:

Khodjanyazova et al

Abstract:

A model of cyclic recurrence (~ 0. 6 – 1.2 myr) of three fusulinid assemblages in the Middle Pennsylvanian siliciclastic-carbonate succession of the Donets Basin is proposed. Each cycle records progressive turnover of assemblages in shallow marine environments in response to sea-level and regional climate change. A Hemifusulina-assemblage (A), adapted to cooler and reduced salinity seawater records the onset of sea level rise accompanied by humid climatic conditions. Sea level high stand is captured by the Beedeina–Neostaffella–Ozawainella–Taitzehoella (or Beedeina-dominated) assemblage (B), characteristic of relatively deeper-water environments. The B assemblage is successively replaced by the most diverse population of the warm-water Fusulinella-dominated assemblage (C). This assemblage, which occurs in the upper limestones of each fusulinid cycle records the onset of sea level fall accompanied by a shift to drier conditions and likely increased seawater salinity.

The proposed model permits robust interbasinal correlation of the Pennsylvanian successions of the Tethyan realm. Fusulinids of the A and C2 assemblages are the most provincial and therefore the most useful for paleogeographic reconstructions. Specifically, they delineate originally contiguous regions that subsequently were dispersed hundreds to thousands of kilometers, whereas fusulinids of the B assemblage hold the highest potential for global correlation. Extinction at the Moscovian – Kasimovian transition of fusulinid genera of the A and B assemblages, which inhabited predominately cooler and normal salinity (perhaps hyposaline) waters, can be explained by the onset of global warming in the earliest Late Pennsylvanian. Fusulinid assemblages define various types of distribution patterns that differ by tectonic setting of the studied basins suggesting that fusulinid assemblage patterns hold potential for reconstructing the paleogeography and tectonic evolution of Pennsylvanian basins of eastern Laurasia.