How big did Your Abelisaur Grow? (and explaining Stromer's Riddle)

A large abelisaurid (Dinosauria, Theropoda) from Morocco and comments on the Cenomanian theropods from North Africa

Authors:

Chiarenza et al

Abstract:

We describe the partially preserved femur of a large-bodied theropod dinosaur from the Cenomanian “Kem Kem Compound Assemblage” (KKCA) of Morocco. The fossil is housed in the Museo Geologico e Paleontologico “Gaetano Giorgio Gemmellaro” in Palermo (Italy). The specimen is compared with the theropod fossil record from the KKCA and coeval assemblages from North Africa. The combination of a distally reclined head, a not prominent trochanteric shelf, distally placed lesser trochanter of stout, alariform shape, a stocky shaft with the fourth trochanter placed proximally, and rugose muscular insertion areas in the specimen distinguishes it from Carcharodontosaurus, Deltadromeus and Spinosaurus and supports referral to an abelisaurid. The estimated body size for the individual from which this femur was derived is comparable to Carnotaurus and Ekrixinatosaurus (up to 9 meters in length and 2 tons in body mass). This find confirms that abelisaurids had reached their largest body size in the “middle Cretaceous,” and that large abelisaurids coexisted with other giant theropods in Africa. We review the taxonomic status of the theropods from the Cenomanian of North Africa, and provisionally restrict the Linnean binomina Carcharodontosaurus iguidensis and Spinosaurus aegyptiacus to the type specimens. Based on comparisons among the theropod records from the Aptian-Cenomanian of South America and Africa, a partial explanation for the so-called “Stromer’s riddle” (namely, the coexistence of many large predatory dinosaurs in the “middle Cretaceous” record from North Africa) is offered in term of taphonomic artifacts among lineage records that were ecologically and environmentally non-overlapping. Although morphofunctional and stratigraphic evidence supports an ecological segregation between spinosaurids and the other lineages, the co-occurrence of abelisaurids and carcharodontosaurids, two groups showing several craniodental convergences that suggest direct resource competition, remains to be explained.

pop sci link too.

Al Noor Tower Rendering

link.

I see you.

You cannot hide from me.

ahem.

Evidence the First Few Thousand Years After Chicxulub Impact the Climate Cooled

Palynological evidence for prolonged cooling along the Tunisian continental shelf following the K–Pg boundary impact

Authors:

Velekoop et al

Abstract:

The Cretaceous–Palaeogene (K–Pg) boundary mass extinction event is related to major global environmental changes resulting from a large extraterrestrial impact. Organic-walled cyst-producing marine dinoflagellates (dinocysts) survived the K–Pg mass-extinction relatively unscathed, making them ideally suited for reconstructing these environmental changes. So far, one of the best dinocyst records available is from the K–Pg boundary Global Stratotype Section and Point (GSSP) at El Kef (NW Tunisia). There, the dinocyst record shows major fluctuations across the boundary, likely reflecting strong responses to environmental changes. These fluctuations have so far not been confirmed by other studies. Therefore, in this study we performed a high-resolution marine palynological study on a closely spaced sample set from the Elles section, some 75 km south of El Kef, in order to generate a palaeoenvironmental and palaeoclimatic record across the K–Pg boundary to allow verification and refinement of earlier reported environmental changes. To better constrain the reconstructions based on qualitative biotic proxies we employed the quantitative sea surface temperature proxy TEX86. Unfortunately, the TEX86 proxy record of the studied section is compromised because of post-depositional oxidation. However, the diverse dinocyst assemblages at Elles show strong fluctuations similar to the El Kef record, therefore confirming the earlier recorded signals, showing rapid, regionally consistent changes. These records imply that the latest Cretaceous was characterized by a gradual cooling trend and the onset of relative sea level fall. Within the immediate post-extinction interval, in the first thousands of years after the impact, dinocyst assemblages reveal multiple incursions of higher-latitude dinocyst species implying repeated pulses of cooling. These results signify that the earliest Danian climatic and environmental conditions were relatively unstable across the Tunisian shelf.

Aegirocassis benmoulae: a 2+ Meter (!!!) Long, Filter Feeding Anomalocaridid From Tremadocian Ordovician Morocco

Anomalocaridid trunk limb homology revealed by a giant filter-feeder with paired flaps

Authors:

Van Roy et al

Abstract:

Exceptionally preserved fossils from the Palaeozoic era provide crucial insights into arthropod evolution, with recent discoveries bringing phylogeny and character homology into sharp focus. Integral to such studies are anomalocaridids, a clade of stem arthropods whose remarkable morphology illuminates early arthropod relationships and Cambrian ecology. Although recent work has focused on the anomalocaridid head, the nature of their trunk has been debated widely. Here we describe new anomalocaridid specimens from the Early Ordovician Fezouata Biota of Morocco, which not only show well-preserved head appendages providing key ecological data, but also elucidate the nature of anomalocaridid trunk flaps, resolving their homology with arthropod trunk limbs. The new material shows that each trunk segment bears a separate dorsal and ventral pair of flaps, with a series of setal blades attached at the base of the dorsal flaps. Comparisons with other stem lineage arthropods indicate that anomalocaridid ventral flaps are homologous with lobopodous walking limbs and the endopod of the euarthropod biramous limb, whereas the dorsal flaps and associated setal blades are homologous with the flaps of gilled lobopodians (for example, Kerygmachela kierkegaardi, Pambdelurion whittingtoni) and exites of the ‘Cambrian biramous limb’ . This evidence shows that anomalocaridids represent a stage before the fusion of exite and endopod into the ‘Cambrian biramous limb’, confirming their basal placement in the euarthropod stem, rather than in the arthropod crown or with cycloneuralian worms. Unlike other anomalocaridids, the Fezouata taxon combines head appendages convergently adapted for filter-feeding with an unprecedented body length exceeding 2 m, indicating a new direction in the feeding ecology of the clade. The evolution of giant filter-feeding anomalocaridids may reflect the establishment of highly developed planktic ecosystems during the Great Ordovician Biodiversification Event.

First Continental Biota (Inclusive of Cladotherian Mammals!) From Bathonian Jurassic Morocco Found

Guelb el Ahmar (Bathonian, Anoual Syncline, eastern Morocco): First continental flora and fauna including mammals from the Middle Jurassic of Africa

Authors:

Haddoumi et al

Abstract:

We report the discovery in Mesozoic continental “red beds” of Anoual Syncline, Morocco, of the new Guelb el Ahmar (GEA) fossiliferous sites in the Bathonian Anoual Formation. They produced one of the richest continental biotic assemblages from the Jurassic of Gondwana, including plants, invertebrates and vertebrates. Both the sedimentological facies and the biotic assemblage indicate a lacustrine depositional environment. The flora is represented by tree trunks (three families), pollen (13 species, five major clades) and charophytes. It suggests local forests and humid (non-arid) conditions. The vertebrate fauna is dominated by microvertebrates recovered by screening-washing. It is rich and diverse, with at least 29 species of all major groups (osteichthyans, lissamphibians, chelonians, diapsids, mammals), except chondrichthyans. It includes the first mammals discovered in the Middle Jurassic of Arabo-Africa. The GEA sites yielded some of the earliest known representatives of osteoglossiform fishes, albanerpetontid and caudate amphibians, squamates (scincomorphans, anguimorphan), cladotherian mammals, and likely choristoderes. The choristoderes, if confirmed, are the first found in Gondwana, the albanerpetontid and caudatan amphibians are among the very few known in Gondwana, and the anguimorph lizard is the first known from the Mesozoic of Gondwana. Mammals (Amphitheriida, cf. Dryolestida) remain poorly known, but are the earliest cladotherians known in Gondwana.The GEA biotic assemblage is characterized by the presence of Pangean and Laurasian (especially European) taxa, and quasi absence of Gondwanan taxa. The paleobiogeographical analysis suggests either a major fossil bias in Gondwana during the Middle Jurassic, and an overall vicariant Pangean context for the GEA assemblage, or alternatively, noticeable Laurasian (European) affinities and North-South dispersals. The close resemblance between the Bathonian faunas of GEA and Britain is remarkable, even in a Pangean context. The similarity between the local Anoual Syncline Guelb el Ahmar and Ksar Metlili faunas raises questions on the ?Berriasian age of the latter.

Coniacian/Santonian Cretaceous Marine Moroccan Records of the Oceanic Anoxic Event 3

Marine palynology of the Oceanic Anoxic Event 3 (OAE3, Coniacian – Santonian) at Tarfaya, Morocco, NW Africa – transition from preservation to production controlled accumulation of marine organic carbon

Author:

Prauss

Abstract:

The paper presented is the first comprehensive, fully quantitative, high resolution study of marine palynology from an OAE3 black-shale environment. It is based on 175 m core spanning the upper Turonian to lower Santonian at Tarfaya, Morocco, NW Africa, which has been sampled from centimetre to 3 m intervals. The results are integrated and discussed with lithology and geochemistry data to (1) distinguish between potential changes in production and preservation of total organic carbon (TOC) accumulation and (2) constrain the stratigraphic position of the Oceanic Anoxic Event 3 (OAE3).

The succession is characterized by increased total organic carbon (TOC), varying between 1% and 19% (average about 6%). Distinct black-shale horizons of variable thickness appear episodically throughout the succession, with higher frequency in the late Turonian. Higher TOC contents do not strictly correlate to lithologic black-shales or peaks of a specific taxon of organic-walled algae. The palynomorph spectrum is strongly dominated by organic-walled algae, with the ratio of terrigenous sporomorphs to organic-walled algae (t/m index) varying between zero and 0.05 (average 0.01). The dominance of algal organic matter is corroborated by the prevalence of Type I kerogen identified using Rock-Eval pyrolysis. Dinocyst diversity is low, with the absolute taxa number varying from 7 to 27 between single samples. The peridinioid/gonyaulacoid ratio of dinocysts (p/g ratio) shows strong fluctuations, varying between 1 and 283 (average of about 100).

The upper Turonian interval is dominated by Bosedinia spp., a dinocyst taxon formerly described as abundant only in lacustrine sediments from the Oligocene and Miocene of SE Asia. This dominance is episodically modified by the increase of the warm-temperate waters dinocysts fraction, here mainly represented by the genera Alterbidinium, Isabelidinium and Spinidinium. Within the Coniacian-Santonian, black-shale horizons are limited in number and are concentrated within the upper Coniacian to lower Santonian interval. The dinocysts show alternating, prominent peak abundances of Palaeohystrichophora spp. and the warm-temperate water dinocysts fraction, here mainly represented by the genera Trithyrodinium and Chatangiella. However, a final episode of increased proportions of Bosedinia spp. is confined to a 5 m thick black-shale horizon closely spanning the Coniacian-Santonian boundary.

Changes in the ratio of total sulphur to total organic carbon (TS/TOC) reflect fluctuating oxygen contents of bottom waters throughout the late Turonian to Santonian. These are significantly parallelled by the alternation of dinocysts assemblages suggestive of enhanced upwelling and water column stratification respectively, probably reflecting changes in the mode of TOC accumulation. Accordingly, preservation largely prevails during the late Turonian interval and changes towards increased production within the Coniacian-Santonian. However, a final preservation-event is probably represented by the black-shale horizon closely spanning the Coniacian-Santonian boundary (top Dicarinella concavata foraminifera zone), which may reflect an episodic shutdown of a major upwelling cell. It is thus proposed, that the “culmination” of the OAE3 at Tarfaya may represent intermittent preservation of TOC within an otherwise high productivity environment related to a global cooling trend.

Alanqa saharica: an Albanian/Campanian Cretaceous Azhdarchid Pterosaur With an Unusual Jaw Modification

An unusual modification of the jaws in cf. Alanqa, a mid-Cretaceous azhdarchid pterosaur from the Kem Kem beds of Morocco

Authors:

Martill et al

Abstract:

A jaw bone attributed to the mid-Cretaceous azhdarchid pterosaur Alanqa saharica Ibrahim et al., 2010 is described. The new specimen from the Kem Kem beds of south eastern Morocco is unusual for the presence of expanded rostral bones that protrude above the occlusal surface of the jaws. The function of this unusual osteological modification is uncertain, but it is considered likely that it served a role in food processing, or perhaps jaw stability.

Westphalian Pennsylvanian Carboniferous Trackways From Morocco

First Occurrence of Tetrapod Footprints from Westphalian Strata of the Sidi Kassem Basin, Central Morocco

Authors:

Lagnaoui et al

Abstract:

The Sidi Kassem Basin is the only limnic basin of Westphalian age in Morocco. It is built up of 1,250 m of alluvio-fluvial to lacustrine deposits that have so far yielded plant fossils and invertebrate remains only. Recent exploration for fossil tetrapod footprints in floodplain-deposits of the basin revealed a moderately diverse vertebrate ichnofauna composed of tracks assigned to cf. Batrachichnus Woodworth, 1900; cf. Hylopus Dawson, 1882; Dimetropus Romer and Price, 1940; and Notalacerta Butts, 1891. The tracks can be referred to temnospondyl, anamniote reptiliomorph, non-therapsid synapsid (pelycosaur), and captorhinomorph trackmakers. The described ichnoassemblage is important in at least three aspects: (1) It suggests an Early to mid-Pennsylvanian age for the footprint-bearing strata of the study area. (2) It is the oldest association of tetrapod footprints from Africa. (3) It is the first evidence of the relatively rare ichnogenera cf. Hylopus and Notalacerta outside of North America and Europe. Judged from the variety of tetrapod tracks and previously collected floral remains, the Sidi Kassem Basin must have represented a well-established continental ecosystem during Pennsylvanian time. Further exploration for trace and body fossils of Palaeozoic vertebrates in this basin may be important for the reconstruction of early tetrapod evolution.

The Maastrichtian Cretaceous Mosasaurs of Morocco

Mosasaurids (Squamata) from the Maastrichtian Phosphates of Morocco: Biodiversity, palaeobiogeography and palaeoecology based on tooth morphoguilds

Authors:

Bardet et al

Abstract:

Mosasaurid squamates are the most numerically abundant, and taxonomically/ecologically diverse clade of marine amniotes represented in the Maastrichtian Phosphates of Morocco. With few exceptions, they are faunally typical of the Southern Mediterranean Tethys Margin (around palaeolatitude 25°N) and range from the base to the top of the stage. The Moroccan assemblages include at least 7 genera and 10 species representing a broad spectrum of sizes and morphologies that illustrate several ecological trends. Noteworthy is the predominance of Mosasaurinae which are widespread in contemporaneous outcrops worldwide and constitute 80% and 70% of the total genus/species number respectively. In contrast, Halisauromorpha and Russellosaurina (plioplatecarpines) are scarce and tylosaurines are presently unknown. All of the Moroccan mosasaurids exhibit characteristic tooth morphologies and can be placed into resource partitioning morphoguilds indicative of adaptations for piercing, crushing or cutting. Medium to large predators are found to distribute along the ‘Crush’-‘Cut’ axis of the morphoguild projection, and a new ‘Crush-Cut’ guild, previously unrecognised amongst Mesozoic marine amniotes, accommodates several Prognathodon species. Also of importance is the lack of mosasaurids along the ‘Pierce’-‘Crush’ axis, potentially inferring that these ecological niches were occupied by other marine vertebrates such as selachians and plesiosaurs. In addition, the relative abundance of mosasaurids throughout the Maastrichtian series of the Gantour Basin evidences direct ecological competition or predation phenomena.

Spinosaurus was Really a Croc, erm, Parallel Evolution to a Croc

Semiaquatic adaptations in a giant predatory dinosaur

Authors:

Ibrahim et al

Abstract:

We describe adaptations for a semiaquatic lifestyle in the dinosaur Spinosaurus aegyptiacus. These adaptations include retraction of the fleshy nostrils to a position near the mid-region of the skull and an elongate neck and trunk that shift the center of body mass anterior to the knee joint. Unlike terrestrial theropods, the pelvic girdle is downsized, the hind limbs are short, and all of the limb bones are solid without an open medullary cavity, for buoyancy control in water. The short, robust femur with hypertrophied flexor attachment and the low, flat-bottomed pedal claws are consistent with aquatic foot-propelled locomotion. Surface striations and bone microstructure suggest that the dorsal “sail” may have been enveloped in skin that functioned primarily for display on land and in water.

A Cryptic Coral-crinoid "Hanging Garden" From Eifelian/Givetian Devonian Morocco

Cryptic coral-crinoid "hanging gardens" from the Middle Devonian of southern Morocco

Authors:

Jakubowicz et al

Abstract:

An unusual and exceptionally well preserved cryptic community of cnidarians, crinoids, sponges, and microbes developed in a submarine cavity of Middle Devonian age in the Hamar Laghdad area (Morocco). The biota encrusted the cavity roof and grew predominantly in an upside-down position, forming spectacular "hanging gardens." The investigated assemblage differs strikingly from both its Paleozoic and modern analogues; it constitutes one of a very few known examples of fossil cryptic assemblages developed in relatively deep water settings, and is the first report of a cryptic paleoecosystem dominated by rugose corals. The results support the view that during the middle Paleozoic there was no distinct polarization between open-surface and cryptic faunas in deep-water environments, but keen competition for space already existed in Devonian cryptic assemblages. The regional species pool seems to have been the main determinant of the ecological succession and structure of this cryptic community.

A Turonian Cretaceous Baby Tethysaurus (mosasaur) Found in Morocco

A baby mosasauroid (Reptilia, Squamata) from the Turonian of Morocco – Tethysaurus ‘junior’ discovered?

Authors:

Alexandra Houssaye and Nathalie Bardet

Abstract:

New Middle Turonian mosasauroid remains were discovered in the same large-sized nodule that yielded a specimen referred to Tethysaurus nopcsai Bardet et al., 2003, from the Goulmima region (southern Morocco). They comprise isolated, fragmentary cranial elements (skull and mandible) and some vertebrae. Their very small size suggests a juvenile condition, an observation supported also by anatomical (spongious nature of numerous bone parts), micro-anatomical (loose inner spongiosa) and histological (numerous remains of calcified cartilage deep into the centrum; radial vascular canals) data. These bones belonged to a basal mosasauroid that cannot be distinguished from Tethysaurus nopcsai to which taxon we tentatively assign the material.

New Sauropods from the Campanian Cretaceous of Morocco

Additions to the sauropod dinosaur fauna of the Cenomanian (early Late Cretaceous) Kem Kem beds of Morocco: Palaeobiogeographical implications of the mid-Cretaceous African sauropod fossil record

Authors:

1. Philip D. Mannion (a)
2. Paul M. Barrett (b)

Affiliations:

a. Department of Earth Science and Engineering, Imperial College London, South Kensington Campus, London SW7 2AZ, UK

b. Department of Earth Sciences, Natural History Museum, Cromwell Road, London SW7 5BD, UK

Abstract:

Determining the relationships of mid-Late Cretaceous African taxa is central to understanding the timing and resultant palaeobiogeographical patterns of Gondwanan fragmentation. The early Late Cretaceous (Cenomanian) Kem Kem beds of southeastern Morocco preserve a diverse vertebrate fauna, including sauropod dinosaurs. Sauropod material includes the holotype of the rebbachisaurid diplodocoid Rebbachisaurus garasbae and fragmentary remains representing indeterminate titanosauriforms and rebbachisaurids. Here, we describe two new specimens from the Kem Kem beds. A dorsal neural arch with complex internal pneumaticity is tentatively attributed to a somphospondylan titanosauriform. A caudal vertebra possessing several rebbachisaurid synapomorphies is excavated by a large and pervasive lateral pneumatic foramen, a feature undocumented in other rebbachisaurids. However, caudal vertebrae are currently unknown for the sympatric R. garasbae, so this element could be referable to that taxon or a second, previously unknown, rebbachisaurid species. Interestingly, this new caudal vertebra displays a mosaic of features otherwise restricted to limaysaurine or nigersaurine rebbachisaurids, suggesting a placement basal to these clades, which is the position usually recovered for R. garasbae in phylogenetic analyses. A review of the mid-Cretaceous African sauropod fossil record removes the Cretaceous record of dicraeosaurids from Africa, restricting this clade to a single post-Jurassic occurrence in Argentina. All diagnostic sauropod remains can be attributed to titanosauriforms or rebbachisaurids. Whereas rebbachisaurids were seemingly restricted to northwestern Africa and disappeared post-Cenomanian, titanosauriforms were widespread across the African continent and survived until the latest Cretaceous. The development of the mid-Cretaceous Trans-Saharan Seaway might have acted as a dispersal barrier for rebbachisaurids and other vertebrate groups. In contrast, titanosauriforms might have been able to cross this barrier, but it is possible that they were also unable to disperse, and that northwestern African titanosauriforms were not closely related to taxa from the rest of the African continent. New materials and a better understanding of titanosaur interrelationships will be crucial in teasing these scenarios apart.

Sigilmassasaurus from Cenomanian Cretaceous Morocco is Valid and Not a Carcharodontosaur

A reevaluation of Sigilmassasaurus brevicollis (Dinosauria) from the Cretaceous of Morocco

Authors:

1. Bradley McFeeters (a)
2. Michael J. Ryan (a)
3. Sanja Hinic-Frlog (b)
4. Claudia Schröder-Adams (a)

Affiliations:

a. Dept. of Earth Sciences, Carleton University, Ottawa, ON K1S 5B6, Canada.

b. Dept. of Biology, Carleton University, Ottawa, ON K1S 5B6, Canada.

Abstract:

The original hypodigm of the controversial mid-Cretaceous Moroccan dinosaur Sigilmassasaurus brevicollis is redescribed, and the diagnosis of the taxon is revised. Unambiguously referred material is restricted to cervical and anterior dorsal vertebrae sharing apomorphies with the holotype. A newly recognized diagnostic character of Sigilmassasaurus is the absence of anterior and posterior interzygapophyseal laminae of the neural arch, so that the neural spine directly meets the dorsal margin of the neural canal. A phylogenetic analysis supports the inclusion of Sigilmassasaurus in Tetanurae but not in Carcharodontosauridae. Sigilmassasaurus is distinct from all other theropods known from comparable material and is thus retained as a valid taxon.

An Unbalanced Terrestrial Ecology in Cenomanian Cretaceous Morroco?

Unbalanced food web in a Late Cretaceous dinosaur assemblage

Authors:

1. Läng Emilie (a)
2. Boudad Larbi (b)
3. Maio Laszlo (c)
4. Samankassou Elias (c)
5. Tabouelle Jérôme (d)
6. Tong Haiyan (e)
7. Cavin Lionel (a)

Affiliations:

a. Dpt de Géologie et de Paléontologie, Muséum d’Histoire Naturelle de la Ville de Genève, CP 6434, 1211 Geneva 6, (FNS grant 129923)Switzerland

b. G2FS, Faculté des Sciences et Techniques, BP, 509, Boutalamine, Errachidia, Morocco

c. Section des Sciences de la Terre et de l’Environnement, Université de Genève, 1205 Genève, Switzerland

d. Musée d’Elbeuf – La Fabrique des Savoirs-CREA, 76500 Elbeuf-sur-Seine, France

e. Palaeontological Research and Education Centre, Mahasarakham University, Khamriang, Kantarawichai, Mahasarakham, 44150Thailand

Abstract:

The rich assemblage of continental vertebrates from the Cenomanian Kem Kem Beds (Morocco) is one of the best known and most diversified for the mid Cretaceous period (Cenomanian). This assemblage, however, shows apparent ecological oddities, in particular the overabundance of theropod dinosaurs versus plant-eating dinosaurs. Several hypotheses have been proposed to explain this unbalanced ratio, including a peculiar ecosystem, non-systematic collecting, taphonomic factors, stratigraphic uncertainties and/or behavioral aspects of the dinosaur groups concerned. Except the comparison of proportions of taxa between field and shop data (McGowan and Dyke, 2009), the other hypotheses have not been tested so far because of the lack of accurately measured sections and systematically collected field data. Based on new field data, we test the above-mentioned hypotheses. The analysis is focused on the ratio of plant-eating versus carnivorous dinosaurs.

This dataset confirms the unbalanced ratio; moreover, the stratigraphic distribution of fossils is quantitatively not homogeneous and consequently important to avoid time-averaging, i.e. the mixing of fossils of different ages together into a single unit. The origin of the unbalanced food web among dinosaurs is related neither to non-systematic collecting, nor to stratigraphic biases. The palaeoenvironment seems to be the only likely factor to explain the significantly high proportion of carnivorous versus plant-eating dinosaurs. Indeed, the deltaic palaeoenvironment offered unfavourable conditions for the setting of stable terrestrial vegetation but favoring aquatic life. This aquatic life formed the basic level of an aquatic or semi-aquatic food web, which directly fed top predators, such as theropods in general and spinosaurs in particular.

New Very Large Ordovician Anomalocaridid Discovered

aleontologists have discovered that a group of remarkable ancient sea creatures existed for much longer and grew to much larger sizes than previously thought, thanks to extraordinarily well-preserved fossils discovered in Morocco.

The creatures, known as anomalocaridids, were already thought to be the largest animals of the Cambrian period, known for the "Cambrian Explosion" that saw the sudden appearance of all the major animal groups and the establishment of complex ecosystems about 540 to 500 million years ago. Fossils from this period suggested these marine predators grew to be about two feet long. Until now, scientists also thought these strange invertebrates—which had long spiny head limbs presumably used to snag worms and other prey, and a circlet of plates around the mouth—died out at the end of the Cambrian.

Now a team led by former Yale researcher Peter Van Roy (now at Ghent University in Belgium) and Derek Briggs, director of the Yale Peabody Museum of Natural History, has discovered a giant fossilized anomalocaridid that measures one meter (more than three feet) in length. The anomalocaridid fossils reveal a series of blade like filaments in each segment across the animal's back, which scientists think might have functioned as gills.

In addition, the creature dates back to the Ordovician period, a time of intense biodiversification that followed the Cambrian, meaning these animals existed for 30 million years longer than previously realized.

"The anomalocaridids are one of the most iconic groups of Cambrian animals," Briggs said. "These giant invertebrate predators and scavengers have come to symbolize the unfamiliar morphologies displayed by organisms that branched off early from lineages leading to modern marine animals, and then went extinct. Now we know that they died out much more recently than we thought."

Note: There is a Devonian anomalocaridid already, Schinderhannes bartelsi, which already extended the clades' existence furter than the Ordovician already. This one IS the biggest by far though.