Ankylosaurs, like probably most other dinosaurs, were landlubbing, terrestrial animals without obvious aquatic adaptations. And yet, surprisingly, their fossils are found in marine sedimentary environments more often than most other dinosaurs (except hadrosaurs). Some, like Aletopelta, wound up in shallow or lagoonal environments - Aletopelta's carcass became a reef! - but some, like the Suncor nodosaurid, wound up far away from shore.
New ankylosaurian material from the Upper Cretaceous of Transylvania
Authors:
Ősi et al
Abstract:
Ankylosaurian remains from the Transylvanian Basin, Romania, are extremely rare. More than 100 years after the discovery of the first and only better-known assemblage, namely the type material of Struthiosaurus transylvanicus, new ankylosaurian material has been discovered in the Maastrichtian of the Haţeg Basin, as well as at another locality (Vurpăr), in the Transylvanian Basin, that is described here. The material consists of one tooth in a small jaw fragment (from the Haţeg Basin) and at least two accummulations of associated, as well as several isolated, postcranial elements (from Vurpăr). No diagnostic elements are preserved that would overlap with the type of S. transylvanicus, so we cannot assign any of the new specimens to this species. The tooth shows marked differences compared to those of other anklyosaurs including S. austriacus and Hungarosaurus in having only six, more or less equally sized, apically pointed cusps separated by deep grooves. The postcranial material from Vurpăr represents at least three different individuals. The humerus is the most diagnostic element among the postcranial remains being most similar both in size and morphology to humeri referred to as Struthiosaurus from different European localities, thus here we refer the humerus and probably associated elements preserved in one assemblage to as cf. Struthiosaurus sp.; the remaining specimens from Vurpăr are retained as Nodosauridae indet. Histological studies have confirmed the adult nature of all sampled bones in the Vurpăr ankylosaur material suggesting that these fully grown animals were of similar size to Struthiosaurus, a small-bodied nodosaurid the ontogenetic status of which, however, has never been investigated histologically. The obviously diminished body size of the Transylvanian ankylosaurs compared to other members of the clade could be explained by insular dwarfism using the same histology-based argument as presented for Magyarosaurus.
Partial skull and endocranial cast of the ankylosaurian dinosaur Hungarosaurus from the Late Cretaceous of Hungary: implications for locomotion
Authors:
Ősi et al
Abstract:
A partial skull of ankylosaur from the Upper Cretaceous (Santonian) Csehbánya Formation in Iharkút and the endocranial cast taken from it are described. The morphology of the exoccipital, the elongated 'neck' region of the basioccipital, the shape of the occipital condyle, and the different flexure of the medulla relative to the forebrain unambiguously differentiate this specimen from the basicranium of Struthiosaurus, so it is assigned to Hungarosaurus sp. Whereas the endocranial cast reflects a brain generally similar to those of other ankylosaurs, the dorsally hypertrophied cerebellum (also present is Struthiosaurus transylvanicus) is quite unusual within the group suggesting a more sophisticated cerebral coordination of posture and movement, and perhaps a more cursorial locomotary habit than predicted for other ankylosaurs.
The basal nodosaurid ankylosaur Europelta carbonensis n. gen., n. sp. from the Lower Cretaceous (lower Albian) Escucha Formation of northeastern Spain
Authors:
Kirkland et al
Abstract:
Nodosaurids are poorly known from the Lower Cretaceous of Europe. Two associated ankylosaur skeletons excavated from the lower Albian carbonaceous member of the Escucha Formation near Ariño in northeastern Teruel, Spain reveal nearly all the diagnostic recognized character that define nodosaurid ankylosaurs. These new specimens comprise a new genus and species of nodosaurid ankylosaur and represent the single most complete taxon of ankylosaur from the Cretaceous of Europe. These two specimens were examined and compared to all other known ankylosaurs. Comparisons of these specimens document that Europelta carbonensis n. gen., n. sp. is a nodosaur and is the sister taxon to the Late Cretaceous nodosaurids Anoplosaurus, Hungarosaurus and Struthiosaurus, defining a monophyletic clade of European nodosaurids - the Struthiosaurinae.
An ankylosaurid dinosaur from Mongolia with in situ armour and keratinous scale impressionsAuthors:1. Victoria M. Arbour (a)2. Nicolai L Lech−Hernes (b)3. Tom E Guldberg (c)4. Jørn H Hurum (d)5. Phillip J Currie (a)Affiliations:a. Department of Biological Sciences, CW 405 Biological Sciences Building, University of Alberta, Edmonton, Alberta, Canada, T6G 2E9b. Bayerngas Norge, Postboks 73, N−0216 Oslo, Norwayc. Fossekleiva 9, N−3075 Berger, Norwayd. Natural History Museum (Geology), University of Oslo, Postboks 1172 Blindern, N−0318 Oslo, NorwayAbstract:A Mongolian ankylosaurid specimen identified as Tarchia gigantea is an articulated skeleton including dorsal ribs, the sacrum, a nearly complete caudal series, and in situ osteoderms. The tail is the longest complete tail of any known ankylosaurid. Remarkably, the specimen is also the first Mongolian ankylosaurid that preserves impressions of the keratinous scales overlying the bony osteoderms. This specimen provides new information on the shape, texture, and arrangement of osteoderms. Large flat, keeled osteoderms are found over the pelvis, and osteoderms along the tail include large keeled osteoderms, elongate osteoderms lacking distinct apices, and medium−sized, oval osteoderms. The specimen differs in some respects from other Tarchia gigantea specimens, including the morphology of the neural spines of the tail club handle and several of the largest osteoderms.
Euoplocephalus tutus and the Diversity of Ankylosaurid Dinosaurs in the Late Cretaceous of Alberta, Canada, and Montana, USAAuthors:1. Victoria M. Arbour (a)2. Philip J. Currie (a)Affiliations:a. Department of Biological Sciences, University of Alberta, Edmonton, Alberta, CanadaAbstract:Few ankylosaurs are known from more than a single specimen, but the ankylosaurid Euoplocephalus tutus (from the Late Cretaceous of Alberta, Canada and Montana, USA) is represented by dozens of skulls and partial skeletons, and is therefore an important taxon for understanding intraspecific variation in ankylosaurs. Euoplocephalus is unusual compared to other dinosaurs from the Late Cretaceous of Alberta because it is recognized from the Dinosaur Park, Horseshoe Canyon, and Two Medicine formations. A comprehensive review of material attributed to Euoplocephalus finds support for the resurrection of its purported synonyms Anodontosaurus lambei and Scolosaurus cutleri, and the previously resurrected Dyoplosaurus acutosquameus. Anodontosaurus is found primarily in the Horseshoe Canyon Formation of Alberta and is characterized by ornamentation posterior to the orbits and on the first cervical half ring, and wide, triangular knob osteoderms. Euoplocephalus is primarily found in Megaherbivore Assemblage Zone 1 in the Dinosaur Park Formation of Alberta and is characterized by the absence of ornamentation posterior to the orbits and on the first cervical half ring, and keeled medial osteoderms on the first cervical half ring. Scolosaurus is found primarily in the Two Medicine Formation of Montana (although the holotype is from Dinosaur Provincial Park), and is characterized by long, back-swept squamosal horns, ornamentation posterior to the orbit, and low medial osteoderms on the first cervical half ring; Oohkotokia horneri is morphologically indistinguishable from Scolosaurus cutleri. Dyoplosaurus was previously differentiated from Euoplocephalus sensu lato by the morphology of the pelvis and pes, and these features also differentiate Dyoplosaurus from Anodontosaurus and Scolosaurus; a narrow tail club knob is probably also characteristic for Dyoplosaurus.
A husband and wife team of American paleontologists has discovered a new species of dinosaur that lived 112 million years ago during the early Cretaceous of central Montana.
The new dinosaur, a species of ankylosaur, is documented in the October issue of the Canadian Journal of Earth Sciences. Ankylosaurs are the biological version of an army tank. They are protected by a plate-like armour with two sets of sharp spikes on each side of the head, and a skull so thick that even 'raptors' such as Deinonychus could leave barely more than a scratch.
Bill and Kris Parsons, Research associates of the Buffalo Museum of Science, found much of the skull of the newly described Tatankacephalus cooneyorum resting on the surface of a hillside in 1997. Because the skull was 90% complete, it was possible to justify this fossil as a new species.
"This is the first member of Ankylosauridae to be found within the Early Cretaceous Cloverly Geologic Formation," said Bill Parsons, who characterized the fossil as a transitional evolutionary form between the earlier Jurassic ankylosaurs and the better known Late Cretaceous ankylosaurs.
The skull is heavily protected by two sets of lateral horns, two thick domes at the back, and smaller thickenings around the nasal region. "Heavy ornamentation and horn-like plates would have covered most of the dorsal surface of this dinosaur" said Bill Parsons.
"For years, Bill and Kris have been collecting fossils from a critical time in Earth's history, and their hard work has paid off," said Lawrence Witmer, professor of paleontology at Ohio University who was not involved with this study. "This is a really important find and gives us a clearer view of the evolution of armored dinosaurs. But this is just the first; I'm sure, of what will be a series of important discoveries from this team."
Parsons also illustrated the dermal armour of this new species based on the theory by Museum of the Rockies paleontologist John R. Horner that there was an outer keratinous sheathing on it as found in modern turtle shells and bird beaks. In his new reconstruction, Parsons suggests that Tatankacephalus exhibited complex and colorful patterns rather than the dull appearance suggested in earlier ankylosaur portraits. "According to Horner's theory, many other dinosaurs also had this kind of sheathing and also may have been diversely colored" said Parsons.
As to its name, the broad, short horns on the back of its skull resemble the horns found on a modern buffalo skull and Tatankacephalus loosely translates as 'Buffalo head.' Parsons also noted, "of course any further allusions to the city of Buffalo are completely intentional as well".
